Washington Flora Checklist
A Checklist of the Vascular Plants of Washington State
Hosted by the University of Washington Herbarium


Checklist » Boraginaceae
Boraginaceae   (Borage Family)

Notes: Taxonomy follows recent molecular studies:

Weigend, M., Federico Luebert, Marc Gottschling, Thomas L.P., Couvreur, Hartmut H. Hilger and James S. Miller. 2013. From capsules to nutlets—phylogenetic relationships in the Boraginales. Cladistics (2013) 1–11.

Nazaire, M., and L Hufford. 2012. A Broad Phylogenetic Analysis of Boraginaceae: Implications for the Relationships of Mertensia. Systematic Botany 37(3): pp. 758–783.

Family Synonyms: (none)

Literature: (none)

Family treatment authored by Peter F. Zika; last updated 2/9/2016 by David Giblin
Taxon (native species in boldface) # Synonyms Common Names
Adelinia grande (Douglas ex Lehm.) J.I. Cohen   [Systematic Botany 40(2): 617.] 1 Pacific hound's tongue, grand hound's tongue
Amsinckia lycopsoides Lehm. ex Fisch. & C.A. Mey.   [H&C] 0 tarweed fiddleneck, bugloss fiddleneck
Amsinckia menziesii (Lehm.) A. Nelson & J.F. Macbr. var. intermedia (Fisch & C.A. Mey.) Ganders   [JPM] 2 rancher's fiddleneck, fireweed fiddleneck, Menzie's fiddleneck, small-flowered fiddleneck
Amsinckia menziesii (Lehm.) A. Nelson & J.F. Macbr. var. menziesii    [JPM] 2 rigid fiddleneck, harvest fiddleneck, rancher's fireweed
Amsinckia spectabilis Fisch. & C.A. Mey. var. spectabilis    [JPM] 1 Scouler's fiddleneck, seaside fiddleneck, woolly breeches
Amsinckia tessellata A. Gray var. tessellata    [JPM] 0 tessellate fiddleneck
Anchusa azurea Mill.   [H&C] 0 Italian alkanet, Italian bugloss
Anchusa officinalis L.   [H&C] 0 common alkanet, common bugloss
Asperugo procumbens L.   [H&C] 0 madwort, catchweed
Borago officinalis L.   [H&C] 0 borage, common borage
Buglossoides arvensis (L.) I.M. Johnston   [Draft Revised H&C] 1 field gromwell, corn gromwell
Cryptantha affinis (A. Gray) Greene   [H&C] 1 common cryptantha, slender cryptantha, quill cat's eye
Cryptantha ambigua (A. Gray) Greene   [H&C] 0 wilke's cryptantha, obscure cryptantha, basin cat's eye
Cryptantha celosioides (Eastw.) Payson   [H&C] 3 northern cryptantha, Sheldon's oreocarya, cockscomb oreocarya, buttecandle
Cryptantha circumscissa (Hook. & Arn.) I.M. Johnston   [JPM] 3 matted cryptantha, cushion cryptantha
Cryptantha fendleri (A. Gray) Greene   [H&C] 0 Fendler's cryptantha, sand dune cat's eye
Cryptantha flaccida (Dougl. ex Lehm.) Greene   [H&C] 0 flaccid cryptantha, weakstem cat's eye
Cryptantha gracilis Osterh.   [JPM] 0 narrow-stem cryptantha
Cryptantha grandiflora Rydb.   [Draft Revised H&C; FNA] 1 large-flowered cryptantha
Cryptantha grandiflora Rydb. 0
Cryptantha intermedia (A. Gray) Greene   [H&C] 3 common cryptantha, Clearwater cat's eye
Cryptantha leucophaea (Dougl. ex Lehm.) Payson   [H&C] 1 gray cryptantha, gray cat's eye
Cryptantha pterocarya (Torr.) Greene   [JPM] 1 wingnut cryptantha
Cryptantha rostellata (Greene) Greene   [H&C] 1 beaked cryptantha
Cryptantha scoparia A. Nelson   [H&C] 0 desert cryptantha, pinyon desert cat's eye
Cryptantha simulans Greene   [H&C] 0 pinewoods cryptantha, pinewoods cat's eye
Cryptantha spiculifera (Piper) Payson   [H&C] 2 Snake River cryptantha, Snake River cat's eye
Cryptantha thompsonii I.M. Johnston   [H&C] 0 Thompson's cryptantha, Thompson's cat's eye
Cryptantha torreyana (A. Gray) Greene   [JPM] 2 Torrey's cryptantha, Torrey's cat's eye
Cryptantha watsonii (A. Gray) Greene   [H&C] 0 Watson's cryptantha, Watson's cat's eye
Cynoglossum officinale L.   [H&C] 0 hound's tongue, common hound's tongue, gypsy flower
Echium vulgare L.   [H&C] 0 viper's bugloss, common viper's bugloss
Eritrichium argenteum W. Wight   [Draft Revised H&C; Draft FNA15] 3 pale alpine forget-me-not
Gruvelia pusilla A. DC.   [Draft FNA] 1 little gruvelia
Hackelia ciliata (Dougl. ex Lehm.) I.M. Johnston   [H&C] 0 Okanogan stickseed
Hackelia cinerea (Piper) I.M. Johnston   [H&C] 0 gray stickseed
Hackelia deflexa (Wahlenb.) Opiz var. americana (A. Gray) Fernald & I.M. Johnston   [H&C] 2 nodding stickseed
Hackelia diffusa (Lehm.) I.M. Johnston var. arida (Piper) R.L. Carr   [Gentry and Carr, 1976] 1 steppe stickseed, sagebrush stickseed
Hackelia diffusa (Lehm.) I.M. Johnston var. cottonii (Piper) R.L. Carr   [Gentry and Carr, 1976] 0 Cotton's stickseed
Hackelia diffusa (Lehm.) I.M. Johnston var. diffusa    [Gentry and Carr, 1976] 1 diffuse stickseed
Hackelia floribunda (Lehm.) I.M. Johnston   [H&C] 0 manyflowered stickseed
Hackelia hispida (A. Gray) I.M. Johnston var. disjuncta R.L. Carr   [Gentry and Carr, 1976] 0 sagebrush stickseed
Hackelia hispida (Gray) I.M. Johnston var. hispida    [Gentry and Carr, 1976] 0 rough stickseed
Hackelia micrantha (Eastw.) J.L. Gentry   [H&C] 1 meadow forget-me-not, blue stickseed
Hackelia taylorii Harrod, Malmquist & R.L. Carr   [JBRIT: 7(2): 652-657] 0 Taylor's stickseed
Hackelia venusta (Piper) St. John   [H&C] 0 showy stickseed, lesser showy stickseed
Lappula fremontii (Torr.) Howell var. fremontii 0
Lappula redowskii (Hornem.) Greene var. cupulata (A. Gray) M.E. Jones   [JPM, H&C] 1 cupped stickseed
Lappula redowskii (Hornem.) Greene var. redowskii    [H&C] 3 western stickseed
Lappula squarrosa (Retz.) Dumort.   [JPM, 2nd Edition] 2 European stickseed, bristly-fruited tickweed, bristly stickseed, common stickseed, bristly sheepburr
Lithospermum incisum Lehm.   [Draft Revised H&C] 0 golden puccoon
Lithospermum ruderale Dougl. ex Lehm.   [H&C] 0 Columbian puccoon, western gromwell, western stoneseed
Lycopsis arvensis L.   [Draft Revised H&C] 1 small bugloss, annual bugloss
Mertensia ciliata (Torr.) G. Don   [H&C] 0 streamside bluebells
Mertensia longiflora Greene   [H&C] 0 long-flowered lungwort, trumpet bluebells
Mertensia oblongifolia (Nutt.) G. Don var. amoena (A. Nelson) L.O. Williams   [JPM] 0 sagebrush bluebells
Mertensia oblongifolia (Nutt.) G. Don var. oblongifolia    [JPM] 0 leafy lungwort, languid lady
Mertensia paniculata (Aiton) G. Don var. borealis (J.F. Macbr.) L.O. Williams   [JPM] 0 tall lungwort
Mertensia paniculata (Aiton) G. Don var. paniculata    [JPM] 0 panicled lungwort
Mertensia platyphylla Heller   [H&C] 2 broadleaved bluebells, western bluebells
Mertensia umbratilis Greenm. 0 shade lungwort, shade bluebells
Myosotis arvensis (L.) Hill   [H&C] 0 field scorpiongrass, mouse ear, rough forget-me-not
Myosotis discolor Pers.   [H&C] 1 yellow and blue scorpiongrass, yellow scorpiongrass
Myosotis laxa Lehm.   [H&C] 0 small forget-me-not, bay forget-me-not
Myosotis scorpioides L.   [H&C] 1 common forget-me-not, water forget-me-not, true forget-me-not
Myosotis stricta Link ex Roemer & J.A. Schultes   [ILBC2] 1 blue scorpiongrass
Myosotis sylvatica Hoffm.   [Stace, 1997] 0 wood forget-me-not
Myosotis verna Nutt.   [H&C] 2 spring forget-me-not, spring scorpiongrass, early scorpiongrass, white scorpiongrass
Pectocarya penicillata (Hook. & Arn.) A. DC.   [IFBC] 1 winged pectocarya, shortleaf combseed
Pectocarya setosa A. Gray   [H&C] 0 bristly combseed
Pentaglottis sempervirens (L.) Tausch ex L.H. Bailey   [IFBC] 1 green alkanet
Plagiobothrys bracteatus (Howell) I.M.Johnst.    [Draf Revised H&C] 0 bracted popcorn-flower
Plagiobothrys figuratus (Piper) I.M. Johnston ex M. Peck ssp. figuratus    [Chambers, 1989] 2 fragrant popcorn flower
Plagiobothrys leptocladus (Greene) I.M. Johnston   [H&C] 0 slender branched popcorn flower, alkali popcorn flower
Plagiobothrys nothofulvus (A. Gray) A. Gray   [H&C] 0 rusty popcorn flower
Plagiobothrys scouleri (Hook. & Arn.) I.M. Johnston var. penicillatus (Greene) Cronquist   [H&C] 8 matted popcorn flower
Plagiobothrys scouleri (Hook. & Arn.) I.M. Johnston var. scouleri    [H&C] 3 Scouler's popcorn flower
Plagiobothrys stipitatus (Greene) I.M. Johnston   [JPM] 0 stipitate popcorn flower
Plagiobothrys tenellus (Nutt. ex Hook.) A. Gray   [H&C] 1 slender popcorn flower, Pacific popcorn flower
Pulmonaria ovalis Bast. 1 oval lungwort
Symphytum asperum Lepech.   [H&C] 1 rough comfrey, prickly comfrey
Symphytum grandiflorum DC.   [Stace, 1997] 1 creeping comfrey
Symphytum officinale L.   [H&C] 0 common comfrey
Symphytum orientale L.   [Stace, 1997] 0 white comfrey
Symphytum ×uplandicum Nyman   [Stace, 1997] 0 hybrid comfrey

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