Voucher:WTU (view specimen records)
Notes: A black-fruited species differing from C. douglasii by its thorns, leaf venation, fruit shape, and stamen number (Phipps & O'Kennon 2002). Recognition at species rank prompted considerable research (Brunsfield & Johnson 1990; Dickinson et al. 1996; Love 1999; Dickinson & Love 1997). Holmgren (1997) treats Crataegus punctata Jacq. var. brevispina Douglas ex Hook. as a synonym of C. douglasii, but the syntypes (K) may belong with C. suksdorfii. The situation needs study and leptotypification. Hybrids with C. monogyna are known from Clark Co., WA, and are occasional in the southern Willamette Valley (Love & Feigen 1978).
» Brunsfield, S. J. & F. D. Johnson. 1990. Cytological, morphological and phenological support for specific status of Crataegus suksdorfii (Rosaceae). Madroņo 37: 274-282.
» Dickinson, T. A., S. Belaoussoff, R. M. Love and M. Muniyamma. 1996. North American black-fruited hawthorns. I. Variation in floral construction, breeding sytem correlates and their possible evolutionary significance in Crataegus sect. Douglasii Loudon Folia Geobot. Phytotax. 31: 355-371.
» Dickinson, T. A. and R. M. Love. 1997. What is Douglas hawthorn? Pp. 162-171, In: Kaye, T. N., A. Liston, R. M. Love, D. L. Luoma, R. J. Meinke, and M. V. Wilson, editors. Conservation and Management of Native Flora and Fungi. Native Plant Society of Oregon, Corvallis, OR,
» Love, R. 1999. The western black-fruited hawthornes: more complexity than formerly suspected. Menziesia 4(4): 10-11.
» Love, R. and M. Feingen. 1978. Interspecific hybridization between native and naturalized Crataegus (Rosaceae) in western Oregon. Madroņo 25: 211-217.
» Holmgren, N. H. 1997. Rosaceae, the rose family, pp. 64-158. In: Cronquist, A., A. H. Holmgren, et. al. 1997. Intermountain Flora Volume 3, Part A: Subclass Rosidae (except Fabales). New York Botanical Garden Press. 456 p.
» Phipps, J. B. and R. J. O'Kennon. 2002. New taxa of Crataegus (Rosaceae) from the northern Okanagan-southwestern Shuswap diversity center. Sida 20: 115-144.
Photo © G. D. Carr
(from WTU Image Collection)