Publication: Icon. Filic. 1: 56. 1829.
selected vouchers: WTU
Notes: FNA2: "The distinctive Polypodium scouleri has occasionally been assigned to the genus Goniophlebium because of its anastomosing venation and conspicuous areoles. Its venation pattern can be quite variable, however, and cannot be used as the sole feature distinguishing P . scouleri from P . californicum . Combining venation characteristics with others provided in the key distinguishes it clearly from its congeners in Polypodium . Some evidence suggests that P . scouleri hybridizes with P . californicum (S. A. Whitmore, unpubl.). I. Manton (1951) reported diploid and triploid cytotypes for P . scouleri , and variation in spore size suggests that the species may also include tetraploid populations."