Publication: Pteridologia. 1: 50.
selected vouchers: WTU
Notes: FNA2: "Polystichum imbricans subsp. imbricans grows in the Coast Ranges and the Sierra-Cascade axis. It is isolated in the Wallowa Mountains of eastern Oregon.
Sun forms of Polystichum munitum are often mistaken for P . imbricans ; characteristics of the distal petiolar scales and indusial margins are more reliable than gross morphologic features for distinguishing them. Polystichum imbricans has narrow distal petiolar scales that fall off early; P . munitum has wide distal petiolar scales (the largest more than 1 mm wide) that are persistent. Polystichum imbricans hybridizes readily with P . munitum , the hybrids usually being sterile but in some places forming hybrid swarms because of partial fertility of the hybrids (D. H. Wagner 1979). The hybrids with P . californicum are discussed under that species."