Dryopteris carthusiana
spinulose fern, toothed wood fern

Distribution: Occurring on both sides of the Cascades crest in Washington; Alaska to Oregon, east across the northern U.S. and Canada to eastern North America.

Habitat: Moist woods and streambanks.

Origin: Native

Conservation Status: Not of concern


Perennial, dying back each winter, stems creeping to erect, lacking stolons.


Leaves monomorphic, 15-75 cm long and 10-30 cm broad; petiole 1/4-1/3 length of leaf; scales found at least at base, scattered, tan; blade light green, ovate-lanceolate, 2-3-pinnate-pinnafitid, not glandular; pinnae mostly in same plane as blade, lanceolate-oblong, basal pinnae lanceolate-deltate and slightly reduced, basal pinnules usually longer than others, basal basiscopic pinnule longer than basal acroscopic pinnule, pinnule margins serrate with spiny teeth.


Indusia glands absent, round-reniform, attached at narrow sinus; spores brownish.

Accepted Name:
Dryopteris carthusiana (Vill.) H.P. Fuchs
Publication: Bull. Soc. Bot. France. 105: 339. 1959. 1959.

Synonyms & Misapplications:
Dryopteris austriaca (Jacq.) Schinz & Thell. var. spinulosa (O.F. Muell.) Fiori
Dryopteris spinulosa (O.F. Muell.) Watt
Polypodium carthusianum Vill.
Polypodium spinulosum O.F. Muell.
Additional Resources:

PNW Herbaria: Specimen records of Dryopteris carthusiana in the Consortium of Pacific Northwest Herbaria database.

WA Flora Checklist: Dryopteris carthusiana checklist entry.

E-Flora BC: Dryopteris carthusiana atlas page.

CalPhotos: Dryopteris carthusiana photos.

USDA Plants: Dryopteris carthusiana information.

24 photographs:
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